Charline Smadi


I am interested in Probability Theory applied to Ecology and Evolution.
Research interests: branching processes, random environment, eco-evolution, varying size population models, selective sweeps, recombinations, clonal interference, mate choice.



  1. M. Voinson, A. Alvergne, S. Billiard, C. Smadi, Stochastic dynamics of an epidemic with recurrent spillovers from an endemic reservoir, Journal of Theoretical Biology, 457:37-50, 2018.
  2. C. Smadi, H. Leman, V. Llaurens, Looking for the right mate in diploid species: how does genetic dominance affect spatial differentiation in a sexual trait?, Journal of Theoretical Biology, 447:154-170, 2018.
  3. C. Coron, M. Costa, H. Leman, C. Smadi, A stochastic model for speciation by mating preferences, Journal of Mathematical Biology, 76:1421–1463, 2018.
  4. C. Smadi, The effect of recurrent mutations on genetic diversity in a large population of varying size, Acta Applicandae Mathematicae, 149(1): 11-51, 2017 (arXiv:1511.03538).
  5. S. Billiard, C. Smadi, The interplay of two mutations in a population of varying size: a stochastic eco-evolutionary model for clonal interference, Stochastic Processes and their Applications, 127(3): 701-748, 2017 (arXiv:1505.04550).
  6. R. Brink-Spalink, C. Smadi, Genealogies of two linked neutral loci after a selective sweep in a large population of varying size, Advances in Applied Probability, 49(1): 279-326, 2017 (arXiv:1502.03837).
  7. S. Palau, J.C. Pardo, C. Smadi, Asymptotic behaviour of exponential functionals of Lévy processes with applications to random processes in random environment, ALEA, Lat. Am. J. Probab. Math. Stat., 13, 1235–1258, 2016.
  8. C. Smadi, V.A. Vatutin, Reduced two-type decomposable critical branching processes with possibly infinite variance, Markov Processes and Related Fields, 21(2): 311-358, 2016 (arXiv:1508.06653).
  9. C. Smadi, An Eco-Evolutionary approach of Adaptation and Recombination in a large population of varying size, Stochastic Processes and their Applications, 125(5): 2054 – 2095, 2015, (arXiv:1402.4104).
  10. V. Bansaye, J.C. Pardo, C. Smadi, On the extinction of Continuous State Branching Processes with catastrophes, Electronic Journal of Probability, 18 no. 106, 1-31, 2013.


  1. A. Gonzalez Casanova, C. Smadi, Multidimensional Lambda-Wright-Fisher processes with general frequency-dependent selection (arXiv:1903.06406).
  2. V. Bansaye, J.C. Pardo, C. Smadi, Extinction rate of continuous state branching processes in critical Lévy environments, (arXiv:1903.06058).
  3. C. Coron, M. Costa, F. Laroche, H. Leman, C. Smadi, Emergence of homogamy in a two-loci stochastic population model, (arXiv:1902.07926).
  4. F. Lavallée, C. Smadi, I. Alvarez, B. Reineking, F-M. Martin, F. Dommanget, S. Martin, A stochastic individual based model for the growth of a stand of Japanese knotweed including mowing as a management technique, (arXiv:1902.06971).
  5. T. Cordonnier, C. Smadi, G. Kunstler, B. Courbaud, Asymmetric competition, ontogenetic growth and size inequality drive the difference in productivity between two-strata and one-stratum forest stands
  6. A. Bovier, L. Coquille, C. Smadi, Crossing a fitness valley as a metastable transition in a stochastic population model, (arXiv:1801.06473).
  7. S. Billiard, C. Smadi, Stochastic dynamics of three competing clones: Conditions and times for invasion, coexistence and fixation, (arXiv:1607.04656).


Isabelle Alvarez, Alexandra Alvergne, Vincent Bansaye, Sylvain Billiard, Anton Bovier, Loren Coquille, Thomas Cordonnier, Camille Coron, Manon Costa, Benoît Courbaud, Fanny Dommanget, Adrian Gonzalez Casanova, Georges Kunstler, Fabien Laroche, Hélène Leman, Violaine Llaurens, François-Marie Martin, Sophie Martin, Sandra Palau, Juan Carlos Pardo, Björn Reineking, Vladimir Vatutin